"Spatial-specific action of serotonin within the leech midbody ganglion" (2008) Calviño, M.A.; Szczupak, L. Journal of Comparative Physiology A: Neuroethology, Sensory, Neural, and Behavioral Physiology. 194(6):523-531

"Calcium spikes in a leech nonspiking neuron" (2009) Rela, L.; Yang, S.M.; Szczupak, L. Journal of Comparative Physiology A: Neuroethology, Sensory, Neural, and Behavioral Physiology. 195(2):139-150

"Premotor nonspiking neurons regulate coupling among motoneurons that innervate overlapping muscle fiber population" (2009) Rodriguez, M.J.; Perez-Etchegoyen, C.B.; Szczupak, L. Journal of Comparative Physiology A: Neuroethology, Sensory, Neural, and Behavioral Physiology. 195(5):491-500

"The activity of leech motoneurons during motor patterns is regulated by intrinsic properties and synaptic inputs" (2012) Perez-Etchegoyen, C.B.; Alvarez, R.J.; Rodriguez, M.J.; Szczupak, L. Journal of Comparative Physiology A: Neuroethology, Sensory, Neural, and Behavioral Physiology. 198(3):239-251