"The activity of leech motoneurons during motor patterns is regulated by intrinsic properties and synaptic inputs" (2012) Perez-Etchegoyen, C.B.; Alvarez, R.J.; Rodriguez, M.J.; Szczupak, L. Journal of Comparative Physiology A: Neuroethology, Sensory, Neural, and Behavioral Physiology. 198(3):239-251

"Effect of a nonspiking neuron on motor patterns of the leech" (2012) Rodriguez, M.J.; Alvarez, R.J.; Szczupak, L. Journal of Neurophysiology. 107(7):1917-1924