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Referencias:

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• Hanski, I., Cambefort, Y., (1991) Dung beetle ecology., , Princeton University Press,Princeton, NJ
• Holter, P., Scholtz, C.H., Stenseng, L., Desert detritivory: Nutritional ecology of a dung beetle (pachysoma glentoni) subsisting on plant litter in arid south african sand dunes (2009) Journal of Arid Environments, 73, pp. 1090-1094
• Howden, H.F., Ritcher, P.O., The biology of deltochilum gibbosum (fab.) with a description of the larva (1952) The Coleopterists Bulletin, 6 (4), pp. 53-57
• Laza, J.H., Dung-beetle fossil brood balls: The ichnogenera Coprinisphaera Sauer and Quirogaichnus (Coprinisphaeridae) (2006) Ichnos an International Journal of Plant and Animal, 13 (4), pp. 217-235. , DOI 10.1080/10420940600843641, PII P11G38225191VRP8
• Monteith, G.B., Storey, R.I., The biology of cephalodesmius, a genus of dung beetles which synthesizes dung from plant material (coleoptera: Scarabaeidae: Scarabaeinae) (1981) Memoirs of the Queensland Museum, 20, pp. 253-257
• Nunes, R.V., Vaz-de-Mello, F.Z., New brachypterous species of dichotomius hope, with taxonomic notes in the subgenus luederwaldtinia martínez (coleoptera: Scarabaeidae: Scarabaeinae) (2013) Zootaxa, 3609 (4), pp. 411-420
• Philips, T.K., Bell, K.L., Attavicinus, a new generic name for the myrmecophilous dung beetle Liatongus monstrosus (Scarabaeidae: Scarabaeinae) (2008) Coleopterists Bulletin, 62 (1), pp. 67-81. , DOI 10.1649/984.1
• Sanchez, M.V., Genise, J.F., Nest and brood chamber structure of two south American dung beetles: Gromphas lacordairei Brullé and Ontherus sulcator (Fabricius) (Coleoptera: Scarabaeidae: Scarabaeinae) (2008) Coleopterists Bulletin, 62 (1), pp. 49-61. , DOI 10.1649/986.1
• Sánchez, M.V., Krause, J.M., González, M.G., Dinghi, P.A., Genise, J.F., The pupation chamber of dung beetles (coleoptera:Scarabaeidae: Scarabaeinae) The Coleopterists Bulletin, 64 (3), pp. 277-284
• Sánchez, M.V., Laza, J.H., Bellosi, E.S., Genise, J.F., Ichnostratigraphy of middle cenozoic dung beetle brood balls (coprinisphaera) from central patagonia: Insights into the appearance and evolution of the south american grass dominated habitats (2010) Palaeogeography,Palaeoclimatology, Palaeoecology, 297, pp. 633-648
• Scholtz, C.H., Chown, S.L., The evolution of habitat use and diet in the scarabaeoidea: A phylogenetic approach (1995) Biology, Phylogeny and Classification of Coleoptera. Papers celebrating the 80th birthday of Roy A. Crowson. (J. Pakaluk and A. Slipinski, editors)., pp. 355-374. , Museum i Instytut Zoologii PAN, Warsaw, Poland
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• Scholtz, C.H., Harrison, J.Du.G., Grebennikov, V.V., Dung beetle (Scarabaeus (Pachysoma)) biology and immature stages: Reversal to ancestral states under desert conditions (Coleoptera: Scarabaeidae)? (2004) Biological Journal of the Linnean Society, 83 (4), pp. 453-460. , DOI 10.1111/j.1095-8312.2004.00389.x
• (2012) Climatología, Iguazú., , www.smn.gov.ar, Servicio Meteorológico Nacional. Available from:(Accessed April 2013)

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